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- Principal Research Associate
- Genetics, Evolution & Environment
- Div of Biosciences
- Faculty of Life Sciences
- Emeritus Professor of Human Genetics
- Genetics, Evolution & Environment
- Div of Biosciences
- Faculty of Life Sciences


Why is there so much genetic diversity? In spite of the huge amounts of polymorphism found in most populations, in most cases we have no idea. I am interested in the relationship between habitat structure and visible variation in snail shell colour and banding pattern. There is a clear fit between the two, and experiments with a paint that fades at a known rate when exposed to the sun shows that animals of different phenotype choose to spend more, or less, time in sunny patches of the habitat. I have extended this work to natural populations of fruit flies, using eye mutants whose appearance depends on the temperature experienced during development, and am now involved in a Europe-wide survey of snail populations to see if there has been any change in gene frequency over the past fifty years and more in response to the changes in climate that have taken place. I have recently been looking at genetic differences in pain reception in populations from different climates and exposure to thermal stress.
I organise and give all the lectures in the large Genes to Organisms course B1005 and about half those on the course Biology B1008 Environment and Evolution together with a variety of single lectures in other courses. In addition I give large numbers of lectures and presentation to schools and other outside organisations and estimate that I have spoken directly to well over 200,000 schoolchildren during my career. I appear regularly in the press, radio and television talking about genetics, evolution and related issues to the general public.
1971 | Doctor of Philosophy | University of Edinburgh | |
1966 | Bachelor of Science (Honours) | University of Edinburgh |